The correct option is this: ANCESTRAL STATE RECONSTRUCTION SUGGESTING THAT THESE ANIMALS HAD OPSINS THAT WERE SHIFTED TOWARD THE RED.
Ancestral reconstruction is the extrapolation back in time from measured characteristics of individuals to their common ancestors. Opsins are a group of light sensitive proteins that are located in the photoreceptor cells of the retina of the eyes. Having opsin that were shifted toward the red colour will enable the animal to see very well in the night time, thus, the animal will be active during that period.