Which of the above are necessary for increasing the volume of a physoclistous gas bladder? (Select all that apply.)

a. The rete mirabile (a "countercurrent" system of capillaries)
b. The gas gland
c. The oval
d. Lactic acid and CO2 released into the blood of the rete mirabile
e. A net disassociation of O2 from hemoglobin in the rete mirabile due to the Root Effect